From Wikipedia, the free encyclopedia
The red panda (Ailurus fulgens), also called the lesser panda, the red bear-cat, and the red cat-bear, is a
native to the eastern
and southwestern . It has reddish-brown fur, a long, shaggy tail, and a waddling gait due to its shorter front legs, and is slightly larger than a . It is , feeds mainly on , but also eats eggs, , and . It is a , mainly , and is largely
during the day.
The red panda has been classified as
because its wild population is estimated at less than 10,000 mature individuals and continues to decline due to
and , , and , although red pandas are protected by national laws in their range countries.
The red panda is the only living species of the
Ailurus and the
. It has been previously placed in the
families, but the results of
research provide strong support for its
classification in its own family Ailuridae, which along with the ,
families is part of the superfamily . Two
are recognized. It is not closely related to the .
Red panda descending head first
Red panda skull
The head and body length of a red panda measures 50 to 64 cm (20 to 25 in), and its tail is 28 to 59 cm (11 to 23 in). Males weigh 3.7 to 6.2 kg (8.2 to 13.7 lb) and females 3 to 6.0 kg (6.6 to 13.2 lb). They have long, soft, reddish-brown fur on the upper parts, blackish fur on the lower parts, and a light face with tear markings and
features. The light face has white badges similar to those of a raccoon, but each individual can have distinctive markings. Their roundish heads have medium-sized upright ears, black noses, and very dark eyes - almost pitch black. Their long bushy tails with six alternating yellowish red transverse
rings provide balance and excellent
against their habitat of moss- and lichen-covered trees. The legs are black and short with thick fur on the soles of the paws. This fur serves as thermal insulation on snow-covered or icy surfaces and conceals scent glands which are also present on the anus.
The red panda is
as a bamboo feeder with strong, curved and sharp semi-retractile claws standing inward for grasping narrow tree branches, leaves, and fruit. Like the giant panda, it has a “false thumb” which is an extension of the wrist bone. When descending a tree head-first, the red panda rotates its ankle to control its descent, one of the few climbing species to do so.
Red panda sleeping
The red panda is
of the , and ranges from the foothills of western
in the east. Its easternmost limit is the
in China. Its range includes southern ,
in , , the northern mountains of , and in south-western China, in the
and the Gongshan Mountains in . It may also live in south-west Tibet and northern , but this has not been documented. Locations with the highest density of red pandas include an area in the Himalayas that has been proposed as having been a refuge for a variety of endemic species in the . The distribution range of the red panda should be considered disjunct, rather than continuous. A disjunct population inhabits the
Plateau of north-eastern India.
During a survey in the 1970s, signs of red pandas were found in Nepal's . Their presence was confirmed in spring 2007 when four red pandas were sighted at elevations ranging from 3,220 to 3,610 m (10,560 to 11,840 ft). The species' westernmost limit is in
located farther west of the Dhorpatan Hunting Reserve. Their presence was confirmed in 2008.
The red panda lives between 2,200 and 4,800 m (7,200 and 15,700 ft) altitude, inhabiting areas of moderate temperature between 10 and 25 °C (50 and 77 °F) with little annual change. It prefers mountainous mixed
forests, especially with old trees and dense understories of .
The red panda population in
is larger and more stable than the Yunnan population, suggesting a southward expansion from Sichuan into Yunnan in the .
The red panda has become
from the Chinese provinces of , , Shaanxi, and .
Distribution of the red panda is disjointed, with two
subspecies:
Western red panda A. f. fulgens (, 1825) lives in the western part of its range, in , , , and .
Styan's red panda A. f. styani lives in the east-north-eastern part of its range, in southern China and northern Burma.
A. f. styani has been described by
in 1902 based on one skull from a specimen collected in Sichuan.
distinguished A. f. styani from A. f. fulgens by its longer winter coat and more abundant blackness in the , bigger skull, more strongly curved forehead, and more robust teeth. His description is based on skulls and skins collected in Sichuan,
close to the border of Yunnan, and Upper Burma.
The Styan's red panda is supposedly larger and darker in color than the Western member of the species, but with considerable variation in both subspecies, and some individuals may be brown or yellowish brown rather than red.
River is often considered the natural division between the two subspecies, where it makes a curve around the eastern end of the Himalayas, although some authors suggest A. f. fulgens extends farther eastward, into China.
Sounds of red panda twittering
it is solitary except during mating season. The species is generally quiet except for some , tweeting, and whistling communication sounds. It has been reported to be both
and , sleeping on tree branches or in tree hollows during the day and increasing its activity in the late afternoon and early evening hours. It sleeps stretched out on a branch with legs dangling when it is hot, and curled up with its tail over the face when it is cold. This panda is very heat sensitive, with an optimal “well-being” temperature between 17 and 25 °C (63 and 77 °F), and cannot tolerate temperatures over 25 °C (77 °F).
Red panda standing
Shortly after waking, red pandas clean their fur like a cat, licking their front paws and then rubbing their backs, torsos, and sides. They also rub their backs and bellies along the sides of trees or rocks. Then they patrol their territories,
and a weak musk-smelling secretion from their anal glands. They search for food running along the ground or through the trees. Red pandas may alternately use their fore paws to bring food to their mouths or place food directly into their mouths.
Predators of the red panda include the ,
(), and humans. If they feel threatened or sense danger, they may try to escape by climbing a rock column or tree. If they can no longer flee, they stand on their hind legs to make themselves appear larger and use the sharp claws on their front paws to defend themselves. The red panda
became a visitor attraction in Japan for his ability to stand upright for ten seconds at a time.
Red panda gnawing
Red pandas are excellent climbers, and forage largely in trees. They eat mostly , and may eat small mammals, birds, eggs, flowers, and berries. In captivity, they were observed to eat birds, flowers,
leaves, and bark and fruits of maple, , and mulberry.
Like the giant panda, they cannot digest , so they must consume a large volume of bamboo to survive. Their diets consist of about two-thirds bamboo, but they also eat mushrooms, roots, acorns, lichens, and grasses. Occasionally, they supplement their diets with fish and insects. They do little more than eat and sleep due to their low-calorie diets.[]
are more easily digested than leaves, exhibiting the highest digestibility in summer and autumn, intermediate digestibility in the spring, and lowest digestibility in the winter. These variations correlate with the nutrient contents in the bamboo. Red pandas process bamboo poorly, especially the cellulose and cell wall components. This implies microbial digestion plays only a minor role in their digestive strategy. To survive on this poor-quality diet, they have to eat the high-quality sections of the bamboo plant, such as the tender leaves and shoots, in large quantities, over 1.5 kg (3.3 lb) of fresh leaves and 4 kg (8.8 lb) of fresh shoots daily. This food passes through the digestive tract fairly rapidly (about 2–4 hr) so as to maximize nutrient intake. Red pandas can taste artificial sweeteners, such as , the only nonprimates known to do so.
Red panda tending its cub
Red pandas are able to reproduce around 18 months of age, and are fully mature at two to three years. Adults rarely interact in the wild except to mate. Both sexes may mate with more than one partner during the mating season from mid-January to early March. A few days before birth, females begin to collect material, such as brushwood, grass, and leaves, to build a nest, which is normally located in a hollow tree or a rock crevice. After a gestation period of 112 to 158 days, the female gives birth in mid-June to late July to one to four blind and deaf cubs weighing 110 to 130 g (3.9 to 4.6 oz) each.
After birth, the mother cleans the cubs and can then recognize each by its smell. At first, she spends 60% to 90% of her time with the cubs. After the first week, the mother starts spending more time outside the nest, returning every few hours to nurse and groom the cubs. She moves the young frequently among several nests, all of which she keeps clean. The cubs start to open their eyes at about 18 days of age. By about 90 days, they have achieved full adult fur and coloring, and begin to venture out of the nest. They also start eating solid foods at this point, weaning at around six to eight months of age. The cubs stay with their mother until the next litter is born in the following summer. Males rarely help raise the young, and only if they live in pairs or in small groups.
Their average lifespan is between eight and 10 years, but individuals have been known to reach 15 years.
Captive red panda
The primary threats to red pandas are direct harvest from the wild, live or dead, competition with domestic livestock resulting in habitat degradation, and
resulting in habitat loss or fragmentation. The relative importance of these factors is different in each region, and is not well understood. For instance, in India, the biggest threat seems to be habitat loss followed by , while in China, the biggest threat seems to be hunting and poaching. A 40% decrease in red panda populations has been reported in China over the last 50 years, and populations in western Himalayan areas are considered to be lower.
Deforestation can inhibit the spread of red pandas and exacerbate the natural population subdivision by
and , leading to severe fragmentation of the remaining wild population. Fewer than 40 animals in four separate groups share resources with humans in Nepal's , where only 6% of 1,710 km2 (660 sq mi) is preferred red panda habitat. Although direct competition for food with domestic livestock is not significant,
can depress bamboo growth by trampling.
Small groups of animals with little opportunity for exchange between them face the risk of inbreeding, decreased genetic diversity, and even extinction. In addition, clear-cutting for firewood or agriculture, including hillside terracing, removes old trees that provide maternal dens and decreases the ability of some species of bamboo to regenerate.
In south-west China, red pandas are hunted for their fur, especially for the highly valued bushy tails from which hats are produced. In these areas, the fur is often used for local cultural ceremonies. In weddings, the bridegroom traditionally carries the hide. The "good-luck charm" red panda-tail hats are also used by local newly-weds. This practice may be quite old, as the red panda seems to be depicted in a 13th-century Chinese pen-and-ink scroll showing a hunting scene. Little or no mention of the red panda is made in the culture and folklore of Nepal.
In the past, red pandas were captured and sold to . Angela Glatston reported she had personally handled 350 red pandas in 17 years.
Due to , this number has decreased substantially in recent years, but poaching continues, and red pandas are often sold to private collectors at exorbitant prices. In some parts of Nepal and India, red pandas are kept as pets.
The red panda has a naturally low birth rate (usually single or twin births per year), and a high death rate in the wild.
Closeup of red panda
Red panda resting on a tree
The red panda is listed in . The species has been classified as
since 2008 because the global population is estimated at about 10,000 individuals, with a decreas only about half of the total area of potential habitat of 142,000 km2 (55,000 sq mi) is actually being used by the species. Due to their shy and secretive nature, and their largely nocturnal habits, observation of red pandas is difficult. Therefore, population figures in the wild are determined by population density estimates and not direct counts.
Worldwide population estimates range from fewer than 2,500 to between 16,000 and 20,000 individuals. In 1999, the total population in China was estimated at between 3,000 and 7,000 individuals. In 2001, the wild population in India was estimated at between 5,000 and 6,000 individuals. Estimates for Nepal indicate only a few hundred individuals. No records from Bhutan or Burma exist.
Reliable population numbers are hard to find, partly because other animals have been mistaken for the red panda. For instance, one report from Burma stated that red pandas were still fairly common in some areas, and was accompanied by a photograph of a "red panda" as proof. The photograph in question depicted a species of .
The red panda is protected in all range countries, and hunting is illegal. Beyond this, conservation efforts are highly variable between countries:
China has 35 protected areas covering about 42.4% of red panda habitat.
India has 20 protected areas with known or possible red panda populations in Sikkim, Arunachal Pradesh, and
such as , , and , and a coordinated conservation policy for the red panda.
In Nepal, known populations occur in , , , , , , and in .
Bhutan has five protected areas that support red panda populations.
Burma has , of which at least one hosts red panda populations.
A community-managed forest in
of eastern Nepal is home to 15 red pandas which generate household income through tourism activities, including home stays. Villagers in the high-altitude areas of Arunachal Pradesh have formed the Pangchen Red Panda Conservation Alliance comprising five villages with a community-conserved forest area of 200 km2 (77 sq mi) at an altitude of 2,500 m (8,200 ft) to over 4,000 m (13,000 ft).
Red panda at Prospect Park Zoo, New York, US
The red panda is quite adaptable to living in captivity, and is common in zoos worldwide. By 1992, more than 300 births had occurred in captivity, and more than 300 individuals lived in 85 institutions worldwide. By
individuals were in North American zoos alone. As of 2006, the international studbook listed more than 800 individuals in zoos and parks around the world. Of these, 511 individuals of subspecies A. f. fulgens were kept in 173 institutions and 306 individuals of subspecies A. f. styani were kept in 81 institutions.
The international studbook is currently managed at the
in the . In cooperation with the International Red Panda Management Group, they coordinate the
in North America, the
in Europe, and other
programs in , India, , and China. In 2009, Sarah Glass, curator of red pandas and special exhibits at the
in , was appointed as coordinator for the North American Red Panda Species Survival Plan. The Knoxville Zoo has the largest number of captive red panda births in the Western Hemisphere (101 as of August 2011). Only the Rotterdam Zoo has had more captive births worldwide.
in , India, successfully released four captive-bred red pandas to the wild in August and November 2003.
Three red panda cubs were born in captivity at
in New Zealand in December 2012, doubling the number held there.
Red panda in a ginkgo tree
The most often cited example of keeping red pandas as pets is by . Pandas were presented to Gandhi's family as a gift and they were then housed in "a special tree house".
Main article:
classification of the red panda has been controversial since it was discovered. French zoologist
initially described the red panda in 1825, and classified it as a close relative of the raccoon (), though he gave it the genus name Ailurus, (from
α?λουρο?, "cat"), based on superficial similarities with domestic cats. The specific epithet is the
adjective fulgens, "shining". At various times, it has been placed in the Procyonidae, , with
(until this family was moved into the Ursidae), and in its own family, the . This uncertainty comes from difficulty in determining whether certain characteristics of Ailurus are phylogenetically conservative or are derived and convergent with species of similar ecological habits.
Red panda gnawing on an exfoliated bamboo bush
Evidence based on the , , , , , and
reflect closer affinities with Procyonidae than Ursidae. However, ecological and
specializations and distinct geographical distribution in relation to modern
support classification in the separate family Ailuridae.
research also places the red panda into its own family, Ailuridae, a part of the broad superfamily
that also includes the , , and
It is not a bear, nor closely related to the giant panda, nor a raccoon, nor a lineage of uncertain affinities. Rather it is a basal lineage of musteloid, with a long history of independence from its closest relatives (skunks, raccoons, and otters/weasels/badgers).
— Flynn et al., Whence the Red Panda, p197
The two subspecies are
and . However, the name Ailurus fulgens refulgens is sometimes incorrectly used for A. f. styani. This stems from a
in his 1874 paper "Recherches pour servir à l'histoire naturelle des mammifères comprenant des considérations sur la classification de ces animaux", making A. f. refulgens a . The most recent edition of Mammal Species of the World still shows the subspecies as A. f. refulgens. This has been corrected in more recent works, including A guide to the Mammals of China and Handbook of the Mammals of the World, Volume 1: Carnivores.
Captive red panda
The red panda is considered a
and only distantly related to the giant panda (Ailuropoda melanoleuca), as it is naturally more closely related to the other members of the superfamily
to which it belongs. The
of both pandas (which also was an ancestor for all living bears,
and seals- and musteloids -, , , ...) can be traced back to the Early
tens of millions of years ago, with a wide distribution across .
of the extinct red panda
have been unearthed from China in the east to
in the west. In 1977, a single tooth of Parailurus was discovered in the
Ringold Formation of . This first North American record is almost identical to European specimens and indicates the immigration of this species from Asia. In 2004, a tooth from a red panda species never before recorded in North America was discovered at the
in . The tooth dates from 4.5–7 million years ago. This species, described as , indicates that a second, more primitive ailurine lineage inhabited North America during the . Cladistic analysis suggests that Parailurus and Ailurus are sister taxa. Additional fossils of Pristinailurus bristoli were discovered at the Gray Fossil Site in 2010 and in 2012. The frequency with which panda fossils are being found at Gray Fossil Site suggests the species played a large role in the overall ecosystem of the area.
The discovery in Spain of the postcranial remains of , a Miocene relative to the red panda, supports a sister-group relationship between red pandas and bears. The discovery suggests the red panda's "false thumb" was an adaptation to arboreal locomotion — independent of the giant panda's adaptation to manipulate bamboo — one of the most dramatic cases of
among vertebrates.
Captive red panda
The first known written record of the red panda occurs in a 13th-century Chinese scroll depicting a hunting scene between hunters and the red panda.
Major General ’s 1821 presentation of an article titled "Description of a new Genus of the Class Mammalia, from the Himalaya Chain of Hills Between Nepaul and the Snowy Mountains" at the
in London is usually regarded as the moment the red panda became a bona fide species in Western science. Hardwicke proposed the name "wha" and explained: "It is frequently discovered by its loud cry or call, resembling the word ‘Wha’, often repeating the same: hence is derived one of the local names by which it is known. It is also called Chitwa." Hardwicke's paper was not published until 1827, by which time Frédéric Cuvier had published his description and a figure. Hardwicke's originally proposed taxonomic name was removed from the 1827 publication of his paper with his permission, and naming credit is now given to Cuvier.
Frédéric Cuvier had received the specimen he described from his brother's stepson, , who had sent it "from the mountains north of India". He was the first to use both the binomial name Ailurus fulgens and the vernacular name panda in his description of the species published in 1825 in Histoire naturelle des mammifères. Ailurus is adopted from the
word α?λουρο? (ailouros), meaning "cat". The specific epithet fulgens is
for "shining, bright". Panda is the French name for the
of peace and travellers, who was called upon before starting a difficult journey. Whether this is the origin of the French vernacular name panda remains uncertain. Later publications claim the name was adopted from a .
described a red panda under the name Ailurus ochraceus, of which Pocock concluded it represents the same
as Ailurus fulgens, since the description of the two agree very closely. He subordinated both types to the Himalayan red panda subspecies Ailurus fulgens fulgens.
Captive red panda
The red panda's local names differ from place to place. The
call it sak nam. In Nepal, the species is called bhalu biralo (bear-cat) and habre. The
people of Nepal and Sikkim call it ye niglva ponva and wah donka. The word w?: in , a small, red bear is called tāwām. In the
region of eastern Nepal, the
know red pandas as kaala, which literally means dark because of th villagers of Tibetan origin call them hoptongar.
Additionally,
lists the vernacular names ye and nigálya ponya (Nepal); thokya and thongwa (); oakdonga or wakdonka and woker (Bhotia); saknam sunam (Lepcha). Nigálya may originate from the
word ?????? ni?ālo or n?gālo meaning a particular kind of small bamboo, namely
intermedia, but also refers to a kind of small leopard, or cat-bear. The word pónya may originate from the Nepali word ????? pajā meaning claw, or ????? pa?jā meaning paw of an animal. Nigálya pónya may translate to bamboo claw or paw.
Nigálya pónya, nyala ponga, and poonya are said to mean eater of bamboo.[] The name panda could originate from panjā.
Drawing and description of red panda in the , 1915
In modern , the red panda is called xi?oxióngmāo (小熊猫/小熊貓, lesser or small panda), or 红熊猫/紅熊貓 (hóngxióngmāo, red panda). In contrast, the giant panda is called dàxióngmāo (大熊猫/大熊貓, giant or big panda), or simply xióngmāo (熊猫/熊貓, panda, literally bear-cat).
In English, the red panda is also called lesser panda, though "red" is generally preferred.[] Many other languages also use red or variations of shining/gold or lesser/small in their names for this species. For instance, червена панда in Bulgarian, panda roux in French, and panda rojo in Spanish all mean red panda. Since at least as far back as 1855, one of its French names has been panda éclatant (shining panda). In , it is called kultapanda (gold panda). Variations of lesser panda occur in
petit panda (small panda),
panda menor (lesser panda),
kleine panda (small panda),
малая панда (malaya panda, "small panda"),
???? (aeki panda, "baby panda"), and
レッサーパンダ (ressā panda, a transliteration of English "lesser panda").
Other names attributed to this species include fire cat, bright panda, and common panda.
The red panda was recognized as the state animal of
in the early 1990s, and was the mascot of the Darjeeling Tea Festival.
In 2005, Babu, a male red panda at
in Birmingham, England, escaped and briefly became a media celebrity, before being recaptured. He was subsequently voted " of the Year", the first animal to receive this honor. Rusty, a male red panda at the
in , similarly attracted media attention when he briefly escaped in 2013.
The name of the
web browser is said to have been derived from a nickname of the red panda.
An anthropomorphic red panda was featured as Master Shifu, the
teacher, in the 2008 film , and its sequels
in 2011 and
in 2016. Some comments on the original film showed a lack of awareness about the red panda in the U.S. at the time it was released. Although most of the reviewers got the species correct, some nevertheless mistook it for a tiny wolf, a rodent, and a lemur. In an interview,
also indicated he did not know much about the animal when he first agreed to voice the character. The red panda
inspired the character of Pabu, the so-called "fire ferret" animal companion (primarily of ), in the U.S. animated TV series .
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Larousse, P. (1866–77) Grand dictionnaire universel du XIXe siècle : fran?ais, historique, géographique, mythologique, bibliographique, littéraire, artistique, scientifique
Larousse et Boyer, Paris
Shrestha, T. K. (2003) . Steven Simpson Books. .
Hale, Austin (ed.) (1973) Clause, sentence, and discourse patterns in selected languages of Nepal 4: Word lists. Summer Institute of Linguistics Publications in Linguistics and Related Fields, 40(4). Norman: Summer Institute of Linguistics of the University of Oklahoma. vii, 314 p.
Yonzon, P.B. (1996) Status of wildlife in the Kanchenjunga region. A reconnaissance study report. WWF Nepal Program, Kathmandu
Turner, R.L.
Turner, R.L.
(1958). . London: Rupert Hart-Davis. p. 48.
Glatston, Angela R. (2010). . . p. 61.  .
Catton, Chris (1990). Pandas. pp. 4–5.  .
. MDBG Chinese-English Dictionary. 2011.
. MDBG Chinese-English Dictionary. 2011.
Gervais, M. Paul (1855).
(in French). 2. L. Curmer. p. 23.
. Government of Sikkim 2010.
. . 24 January .
Bounds, Jon. .
Gabriel, T. (24 June 2013). . New York Times 2013.
Day, P. K. (24 June 2013). . Los Angeles Times 2013.
. Mozilla. Archived from
on April 21, .
. BBC Nature 2014.
Keller, Louise (2008). . <.au. Urban Cinefile 2010.
Boyce, Maree (30 June 2008). . media-culture.org.au. MC Culture 2010.
Pappas, Jim (2 June 2008). . . The Trades 2010.
Symkus, Ed (4 June 2008). . <. Wicked Local Watertown 2010.
Vigil, Delfín (1 June 2008). . <. San Francisco Chronicle 2010.
(28 September 2012).
Glatston, Angela (2007a).
(PDF). Rotterdam Zoo 2009.
Glatston, Angela (2007b).
(PDF). Rotterdam Zoo 2009.
ITIS (USDA Integrated Taxonomic Information System).
IUCN/SSC Mustelid, Viverrid, and Procyonid Specialist Group (1994). A. R. Glatston, ed.
(PDF). Gland, Switzerland: IUCN.   2010.
Slattery, J. P O'Brien, S. J. (1995). "Molecular phylogeny of the red panda (Ailurus fulgens)". The Journal of Heredity. Oxford University Press. 86 (6): 413–22.  .
Mace, G.M. and Balmford, A. (2000). “Patterns and processes in contemporary mammalian extinction.” In Priorities for the Conservation of Mammalian Diversity. Has the Panda had its day?, A. Entwhistle and N. Dunstone (eds). Cambridge University Press, Cambridge. pp. 27–52.
Miyashiro (25 August 2006).
(PDF). New Mexico Tech 2010.
Naish, Darren (3 April 2008). . Tetrapod Zoology 2010.
has media related to:
has information related to:
– The world's only non-profit organization dedicated to red panda conservation.
Ailurus fulgens. Retrieved on .
– UK breeding program. Retrieved on .
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